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Anthomyzidae Temporal range: Eocene–Recent PreꞒ Ꞓ O S D C P T J K Pg N
Anthomyza gracilis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Section:	Schizophora
Subsection:	Acalyptratae
Superfamily:	Opomyzoidea
Family:	Anthomyzidae Czerny, 1903
Genera
See text

Anthomyzidae is small, slender, yellow to black flies with narrow and elongated wings, which may have distinct markings. Some species have greatly reduced wings. Fewer than 100 species are known, mostly from Europe. Although they occur in all major regions, they seem to be most varied in the Holarctic region.

Around 20 diverse genera have been placed in the family. Two, Teratomyza and Teratoptera, are now in the Teratomyzidae, and Cyamops and Stenomicra are in the Stenomicridae. Melanthomyza Malloch from Chile should probably not be retained in the family. The remaining genera are very similar to one another.

Description

For terms see Morphology of Diptera

These are minute to small (1.3-4.5 mm), slender flies. They are yellow (sometimes with dark spots or stripes) to black in colour. The postverticals on the head are small, convergent or parallel, and rarely absent. Two or three pairs of frontal bristles, which curve backward, are present and usually preceded by one or more weaker bristles. Interfrontal bristles are absent or present. Peristomal bristles ("false vibrissae") are present. In the more common Anthomyza and Paranthomyza, the lower side of femur 1 has a well-developed spine in apical third. Wings are usually long and narrow and immaculate (sometimes marked). Some species are brachypterous. The costa has a subcostal break and the subcosta is incomplete.

Biology

Larvae have been reported from decaying dicotyledonous plants, from fungi, and in Europe from leaf sheaths of various grasses and of Typha, Scirpus, and Juncus, from Lipara galls on Phragmites. They may be either phytophagous or saprophagous, but damage to cereals or other plants has not been reported.

Adults are usually found in moist habitats such as damp meadows, marshes, bogs, and damp deciduous or mixed forests with rich undergrowth. Some species inhabit dry grasslands (some species of Anthomyza and the brachypterous Stiphrosoma sabulosum).

Phylogeny and taxa

The family includes a number of genera in two subfamilies.

• Subfamily Anthomyzinae Czerny, 1903
  • Amnonthomyza Roháček, 1993
  • Amygdalops Lamb, 1914
  • Anagnota Becker, 1902
  • Anthomyza Fallén, 1810
  • Apterosepsis Richards, 1962
  • Barbarista Roháček, 1993
  • Carexomyza Roháček, 2009
  • Cercagnota Roháček & Freidberg, 1993
  • Epischnomyia Roháček, 2006
  • Fungomyza Roháček, 1999
  • †Grimalantha Roháček, 1998 (Burdigalian, Dominican amber)
  • Ischnomyia Loew, 1863
  • †Lacrimyza Roháček, 2013 (Lutetian, Baltic Amber)
  • Margdalops Roháček & Barraclough, 2003
  • Mumetopia Melander, 1913
  • Paranthomyza Czerny, 1902
  • Receptrixa Roháček, 2006
  • Reliquantha Roháček, 2014 (Lutetian, Baltic amber)
  • Santhomyza Roháček, 1984
  • Stiphrosoma Czerny, 1928
  • Typhamyza Roháček, 1992
  • Zealantha Roháček, 2007
• Subfamily †Protanthomyzinae Roháček, 1998
  • †Protanthomyza Hennig, 1965 (Lutetian, Baltic Amber, Bitterfeld amber)

References

1. Nello schema di McAlpine, i Clusiidae sono in relazione con il genere Acartophthalmus, che secondo l'analisi cladistica di Buck (2006) va collocato nel clade dei Carnoidea. Vedi Acartophthalmidae.
2. Roháček (2020). "Protanthomyza grimaldii sp. nov., a further member of the extinct subfamily Protanthomyzinae (Diptera, Anthomyzidae) from Baltic amber". ZooKeys (973): 1–15. doi:10.3897/zookeys.973.51435. PMC 7550391. PMID 33110370.

Further reading

• Przemysław Trojan, 1962 Odiniidae, Clusiidae, Anthomyzidae, Opomyzidae, Tethinidae in (series) Klucze do oznaczania owadów Polski, 28,54/58; Muchowki = Diptera, 54/58 Publisher Warszawa : Państwowe Wydawnictwo Naukowe (in Polish)
• Jindřich Roháček, 1996. Revision of Palaearctic Stiphrosoma, including the Anthomyza-laeta group (Diptera, Anthomyzidae). Eur. J. Entomol. 93:89-120, ISSN 1210-5759 European Journal of Entomology
• Jindřich Roháček, 1998. Taxonomic limits, phylogeny and higher classification of Anthomyzidae (Diptera), with special regard to fossil record. Eur. J. Entomol. 95:141-177, ISSN 1210-5759 European Journal of Entomology
• Jindřich Roháček, 2006. A monograph of Palaearctic Anthomyzidae (Diptera), Part 1.published as supplement 1 of the Časopis Slezského zemského muzea, Vol. 55 (2006) 326 pages, 661 black-and-white illustrations. ISSN 1211-3026, ISBN 80-86224-57-0
• Jindřich Roháček, 2007. Zealantha thorpei gen. et sp. nov. (Diptera: Anthomyzidae), first family representative from New Zealand. Zootaxa 1576: 1–13 Zootaxa

Species lists

• West Palaearctic including Russia
• Australasian/Oceanian
• Nearctic
• Japan
• World list

External links

• Encyclopedia of Life - Family Anthomyzidae at EOL
• Diptera.info images

• Anthomyzidae
• Brachycera families